Since there is no connection between the printing heads, they are completely independent in their numbering sequence and if a mishaps occurs affecting one of the 90 printing heads we find to print a complete sheet, it's only this defective head which will put the wrong serial and it can go for quite long before the operator catches the error and repair/readjust (or replace if necessary) the faulty numbering head.
I would believe that each numbering wheel in the printing head can move independently, in part for the operator can easily and rapidly set the serials to the next sequence (between two printing runs). However, I dont know if the stepping is completely mechanical (driven by the rotation of the drum) or electromechanical (driven with electric pulses synchronized with the passes of the sheets)...